Stan Harpole
Postdoctoral Scholar
Website
Education:
Ph.D. Ecology, University of Minnesota
B.S. Botany, University of Washington
Research Interests:
As a community ecologist, I study the mechanisms that control species diversity as a way to predict the consequences of global change to diversity. My work is centered on testing and developing biodiversity theory using experimental and observational studies of terrestrial plant communities in combination with mathematical and statistical models.
Recent Presentations:
Ecological Society of America 2006, Symposium: Multiple Resource Limitation
Recent Publications:
Harpole, W.S. and K.N. Suding. (in press). Frequency-dependence stabilizes competitive interactions among four annual plants. Ecology Letters.
Harpole, W.S., D.L. Potts, K.N. Suding. (in press). Ecosystem responses to water and nitrogen amendment in a California grassland. Global Change Biology.
Elser, J.J., M.E.S. Bracken, E.E. Cleland, D.S. Gruner, W.S. Harpole, H. Hillebrand, J.T. Ngai, E.W. Seabloom, J.B. Shurin, J.E. Smith. (in press). Global analysis of nitrogen and phosphorus limitation of primary producers in freshwater, marine, and terrestrial ecosystems. Ecology Letters.
Fox, J.W., and W.S. Harpole. (in press). Understanding how species loss affects ecosystem function: development and application of a trait-based approach. Ecology.
Harpole, W.S., D. Tilman. 2007. Grassland Species Loss Due to Reduced Niche Dimension. Nature. 446:791-793.
Harpole, W.S., L. Goldstein, R. Aicher. Resource limitation. 2007. In C. D’Antonio, J. Corbin, and M. Stromberg, editors. Ecology and Management of California Grassland. University of California Press. 119-127.
Hillebrand, H., D.S. Gruner, Borer, E.T., M.E. Bracken, E.E. Cleland, J.J. Elser, W.S. Harpole, J.T. Ngai, E.W. Seabloom, J.B. Shurin, J.E. Smith. 2007. Consumer versus resource control of producer diversity depends on ecosystem type and producer community structure. Proceedings of the National Academy of Sciences. 104(26):10904-10909.
Fargione, J., D. Tilman, R. Dybzinski, J. Hille Ris Lambers, C. Clark, W.S. Harpole, J. M. H. Knops, P. B. Reich, M. Loreau. 2007. From selection to complementarity: Shifts in the causes of biodiversity-productivity relationships in a long-term biodiversity experiment. Proceedings of the Royal Society B. 274:871–876.
Harpole, W.S., D. Tilman. 2006. Non-neutral patterns of species abundance and species traits. Ecology Letters. 9:15-23.
Harpole, W.S. 2006. Resource-ratio theory and the control of invasive plants. Plant and Soil. 280:23-27.
Moe, S.J., R.S. Stelzer, M.R. Forman, W.S. Harpole, T. Daufresne, T. Yoshida. 2005. Recent advances in ecological stoichiometry: insights for population and community ecology. Oikos. 109:29-39.
Hille Ris Lambers, J., W.S. Harpole, D. Tilman, J. Knops, P. Reich. 2004. Species and mechanisms contributing to the positive diversity-productivity relationship. Ecology Letters. 7:661-668.
Tilman, G.D., J. Hille Ris Lambers, S. Harpole, R. Dybzinski, J. Fargione, C. Clark, C. Lehman. 2004. Does metabolic theory apply to community ecology? It's a matter of scale. Ecology. 85:1797-1799.
Seabloom, E., W.S. Harpole, O.J. Reichman, D. Tilman. 2003. Resource competition, seed-limitation, and invasion in California grasslands. Proceedings of the National Academy of Sciences. 100:13384-13389.
Parker, I.M., W.S. Harpole, D. Dionne. 1997. Plant community diversity and invasion of the exotic shrub Cytisus scoparius: Testing hypotheses of invasibility and impact. pp. 149-162, in P.V. Dunn and K. Ewing, eds., Ecology and Conservation of the South Puget Sound Prairie Landscape. The Nature Conservancy Press, Seattle.
Publications In Review:Potts, D.L., Harpole, W.S., M.L. Goulden, K.N. Suding. The impact of invasion and subsequent removal of an exotic thistle, Cynara cardunculus, on carbon and water cycling in a coastal California grassland. Biological Invasions.
Favorite Plant: 
Monstera deliciosa